Which of the following scientists discovered HLA genes:

Alexis Carrel (France): Work on vascular suture and the transplantation of blood vessels and organs Peter Brian Medawar: Discovery of acquired immunological tolerance .The graft reaction is an immunity phenomenon. 1950s, induced immunological tolerance to skin allografts in mice by neonatal injection of allogeneic cells. George D. Snell , Jean Dausset: Discoveries concerning genetically determined structures on the cell surface that regulate immunological reactions. H-genes (histocompatibility genes), H-2 gene-mice; Human transplantation antigens (HLA) ----MHC Gerude B. Elion , George H. Hitchings: Discoveries of impoant principles for drug treatment in immunosuppression. Joseph E. Murray: Discoveries concerning organ transplantation in the treatment of human disease. In 1954, the first successful human kidney transplant was performed between twins in Boston. Transplants were possible in unrelated people if drugs were taken to suppress the body's immune reaction. Ref: Kuby Immunology; 6th edition